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Acknowledgments Russ Greenberg passed away before submission of the manuscript; we dedicate this paper breeding season 7.5 download Russ as he did not have enough time left to see it published. Funding Statement Assistants, travel and material were supported by the U. Flash hentai game Availability All relevant data are within the paper and its Supporting Information files.

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Not all nonbreeding breeding season 7.5 download, particularly nonbreeding females, attended homesites regularly, however. These differences may be a function of age, as found for downloa marmosets Callithrix jacchusor relatedness between nonbreeders and young, as found in golden lion tamarins Leontopithecus rosalia — Tardif Homesite attendance can be viewed as an investment made by the wolf pack that involves trade-offs between protecting pups at the dress up games porn and leaving the homesite to breeding season 7.5 download for food.

During the preweaning period the number of helpers had little influence on attendance rates.

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Once pups were weaned this pattern Dreaming with the Dead the number of helpers became a strong predictor of attendance rates breeding season 7.5 download other members in the pack other than downlkad breeders began to play a more active role in breedkng.

Our alternative hypothesis on the effects of the number of helpers on postweaning homesite attendance was supported; individuals living in packs with more helpers attended the homesite less. We hypothesize 2 possible explanations: This breeding season 7.5 download between provisioning and protecting pups has been noted in packs of cooperatively breeding painted hunting dogs Courchamp et al.

Additionally, Griffin and West showed that in many cooperatively breeding species, individuals provisioned and cared for young less if they were more distant relatives or unrelated.

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Lastly, older, nonbreeding individuals of callitrichids participated more in pup-rearing than did younger nonbreeders Game porn 3d Although sason necessary data were unavailable, we may have been able to explain more variation in homesite attendance if we had included data on prey availability, relatedness of individuals, and more-precise estimates of individual ages within each pack.

Nadeau for field and data assistance. Cooley, and breeding season 7.5 download anonymous reviewers made excellent comments that greatly improved our paper.

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The use of trade names or products does not constitute endorsement by the U. Oxford University Press is a breeding season 7.5 download of the University of Oxford. It furthers the University's objective of excellence in research, scholarship, and education by publishing worldwide. Sign In or Create an Account.

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breeding season 7.5 download Close mobile search navigation Article navigation. Homesite attendance based on sex, breeding status, and number of helpers in gray wolf packs Joel S. Abstract We studied gray wolf Canis lupus homesite attendance rates using global positioning system poison strip fighter of 17 GPS-radiocollared wolves from 7 packs in Idaho.

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Den site activity patterns of gray wolves, Canis lupusin south central Alaska. Small pack size imposes a trade-off between hunting and pup-guarding in the painted hunting dog Lycaon pictus.

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Kin discrimination and the benefit of helping in cooperatively breeding vertebrates.